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In 1954, a nuclear bomb is detonated at the moment a giant creature emerges from the ocean. In 1999, Project Monarch scientists Ishiro Serizawa (Ken Watanabe) and Vivienne Graham (Sally Hawkins) investigate a colossal skeleton unearthed in a collapsed mine in the Philippines. They find two giant spores; one dormant and one hatched that made a trail into the sea. In Japan, the Janjira Nuclear Power Plant experiences unusual seismic activity; and Supervisor Joe Brody (Bryan Cranston) sends his wife Sandra (Juliette Binoche) and a team of technicians into the reactor. A tremor breaches the reactor, leaving Sandra and her team unable to escape while the plant collapses.

Fifteen years later, Joe's son Ford (Aaron Taylor-Johnson), a U.S. Navy explosive ordnance disposal officer, returns from a tour of duty to his family in San Francisco but has to immediately depart for Japan after Joe is detained for trespassing in the Janjira quarantine zone. Joe persuades Ford to accompany him to their old home in the quarantine zone to retrieve vital data. They successfully retrieve the data but are discovered by soldiers and taken to a secret facility in the power plant's ruins. After several power failures, a giant winged creature emerges and escapes, destroying the facility. Joe is severely wounded and dies after being evacuated by helicopter to the U.S.S. Saratoga. The incident is reported around the world as an earthquake.


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